behavior/neurological phenotype

• ataxia
  (MGI Ref ID J:142871)

• abnormal motor capabilities/coordination/movement
  (MGI Ref ID J:142871)

• abnormal gait
  (MGI Ref ID J:142871)

• impaired righting response
  (MGI Ref ID J:142871)

mortality/aging

• premature death
  (MGI Ref ID J:142871)

nervous system phenotype

• abnormal hippocampus morphology
  • neurons of the hippocampal gyrus are scattered in irregular wavy layers
  (MGI Ref ID J:142871)

• abnormal cerebral cortex morphology
  • scrambled layering in the cortex
  (MGI Ref ID J:142871)

• abnormal dentate gyrus morphology
  (MGI Ref ID J:142871)

• small cerebellum
  (MGI Ref ID J:142871)

• abnormal cerebellar cortex morphology
  • cerebellum is small with scrambled Purkinje and granule cells
  (MGI Ref ID J:142871)